In order to elucidate the degree of preference in orientations of listeners whether on semantic or prosodic factors under the reconstruction of sense of utterance, an audition experiment was performed based on the material of spontaneous polylogues of native speakers of the Russian and German languages. The conducted perceptive-acoustic experiment proves the hypothesis that the prognostification of the sense of the reconstructed utterance is implemented with the support of semantic and prosodic features on equal grounds. The decision-making between them or their combination is based on verbal variability as an ontological feature of the language and on perceptive-conceptual mnemonic activities of bearers of different cultures.